In simple terms
A friendly intro before the formal notes — no formulas yet.
Energy flows, matter cycles
Sunlight enters an ecosystem, is captured by producers, and is passed along a food chain to consumers — but almost all of it leaks out as heat at every step, so it never comes back. The atoms those organisms are built from behave differently: carbon, nitrogen and the rest are used again and again, released by decomposers and taken up once more by producers.
Picture a relay race run with a leaky bucket of water. The water is the energy: each runner (trophic level) pours what is left into the next runner's bucket, but so much sloshes out on the way that after only a few handovers there is barely a splash left — which is why the race has few legs. The runners' numbered vests are the matter: they are collected at the end, washed, and handed back out for the next race. The water is spent and gone; the vests go round again.
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Producers capture light energy and convert it to chemical energy stored in their biomass (photosynthesis).
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At each feeding step only about 10% of the energy is passed on; the rest is lost — mostly as heat from respiration, some never eaten, some never digested.
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Because so little energy remains, food chains are short and a pyramid of energy is always upright.
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Decomposers break down dead matter and waste, releasing the nutrient atoms so producers can take them up again — matter cycles while energy does not.
Explore the concept
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Full topic notes
Formal explanation with the rigour you need for the exam.
Ecosystems as open systems: energy flows, matter cycles
An ecosystem is an open system: it exchanges both energy and matter with its surroundings. But the two are exchanged in fundamentally different ways. Energy flows through an ecosystem in one direction — it enters (in most ecosystems) as light, is converted to chemical energy by producers, is passed between organisms, and is ultimately lost to the surroundings as heat. It is never recycled, so it must be continually resupplied by the Sun. Matter cycles within an ecosystem. The chemical elements that make up organisms — carbon, nitrogen, phosphorus and the rest — are finite. They pass from the environment into organisms and back again, circulating between the living (biotic) and non-living (abiotic) components, driven above all by decomposers. Holding these two ideas apart — flow versus cycle — is the single most important thing in this topic.
Energy FLOWS: one direction only — in as light, out as heat; lost at every step; never recycled; needs constant resupply from the Sun.
Matter CYCLES: the same finite atoms are used again and again, passing between organisms and the environment.
Open system: an ecosystem exchanges both energy and matter with its surroundings.
The commonest error in this whole topic is saying energy is 'recycled' — it is not.
Trophic levels: producers, consumers and decomposers
A trophic level is an organism's feeding position — how many steps its energy is from the original source. Producers (autotrophs such as plants and algae) form the first level: they make their own organic compounds, usually by photosynthesis, and are the entry point of energy into the ecosystem. Consumers (heterotrophs) obtain energy by feeding on other organisms. A primary consumer (herbivore) eats producers; a secondary consumer eats primary consumers; a tertiary consumer eats secondary consumers. Standing apart from this ladder are the decomposers — chiefly saprotrophic bacteria and fungi — which feed on dead organic matter and waste from every level, breaking it down and releasing inorganic nutrients back to the environment. Decomposers are what make matter cycling possible.
Producers: autotrophs (photosynthesise); first trophic level; the energy entry point.
Consumers: heterotrophs — primary (eat producers), secondary (eat primary), tertiary (eat secondary).
Decomposers: feed on dead matter and waste from all levels; release inorganic nutrients — the key to nutrient cycling.
Producers make organic matter; decomposers break it down — do not confuse the two.
Food chains and food webs
A food chain shows a single sequence of feeding relationships, for example: grass → grasshopper → shrew → owl. The arrows are critical: each arrow points in the direction that energy flows, from the organism being eaten to the organism that eats it (prey → predator). Real ecosystems are far more tangled than a single chain, because most organisms eat, and are eaten by, several species. A food web joins many food chains together to give a realistic map of who feeds on whom. Reading a web, you can trace how energy captured by producers is distributed through the whole community — and see why removing one species can ripple through many others.
Draw and read food-chain arrows as ENERGY arrows: they run from prey to predator (grass → rabbit → fox), never the other way. A surprising number of marks are lost simply by drawing the arrows backwards, as though they meant 'eats'.
Energy transfer and the 10% rule
The transfer of energy from one trophic level to the next is strikingly inefficient. As a rule of thumb, only about 10% of the energy at one level is passed on to the next; roughly 90% is lost. (The real figure varies, commonly between about 5% and 20%, so 10% is an approximation rather than a law.) The energy does not vanish — it goes to three places. Most is lost as heat from respiration: organisms respire much of their food to power movement, growth and, in birds and mammals, keeping warm. On top of that, not all of an organism is eaten (roots, bones, fur and wood are often left), and of what is eaten, not all is digested and absorbed — the indigestible remainder leaves in faeces. Only the small fraction built into new biomass is available to the next level.
Two consequences follow directly. First, food chains are short: after only a few ~90% losses, so little energy remains that no further trophic level can be supported — which is why chains rarely exceed four or five links. Second, a pyramid of energy (the energy, or rate of energy flow in kJ m⁻² yr⁻¹, at each level) is always upright, because each level must hold less than the one below. Pyramids of numbers or biomass can occasionally be inverted — a single tree feeding many insects, for instance — but a pyramid of energy never can.
Respiration (largest loss): energy used for life processes is released as heat and leaves the ecosystem.
Not all eaten: parts of organisms (roots, bones, fur, wood) are never consumed.
Not all digested: of the material eaten, the indigestible part is lost in faeces.
Result: only about 10% is stored as new biomass and passed on — hence the ~90% loss.
The recycling of matter: the carbon cycle
While energy drains away, the atoms are conserved and cycled — and the clearest example is the carbon cycle. Carbon dioxide is removed from the atmosphere by photosynthesis, which fixes it into the organic compounds of producers. That organic carbon then moves along food chains by feeding, as consumers eat other organisms. Carbon returns to the atmosphere as CO₂ through respiration by every organism, and through decomposition, in which decomposers break down dead organisms and waste and respire their carbon. Finally, combustion — the burning of biomass, and of fossil fuels formed from ancient organisms — releases CO₂ as well. Decomposers are pivotal: by releasing carbon and mineral nutrients from dead matter, they return the raw materials of life to the environment so producers can take them up again. This is why matter cycles indefinitely while energy does not.
Photosynthesis: removes CO₂ from the atmosphere, fixing carbon into producer biomass.
Feeding: transfers organic carbon along food chains from one organism to the next.
Respiration: returns CO₂ to the atmosphere from all living organisms.
Decomposition: decomposers break down dead matter and waste, respiring its carbon back to CO₂ and freeing nutrients.
Combustion: burning biomass and fossil fuels releases CO₂ to the atmosphere.
Human impact on energy flow and the carbon cycle
Human activity disturbs these balanced flows — most importantly by moving carbon faster than the cycle can absorb. Burning fossil fuels releases carbon that had been locked away for millions of years, and deforestation both removes photosynthesising producers and, when the timber is burned or left to decompose, adds still more CO₂. Because this input outpaces removal by photosynthesis and the oceans, atmospheric CO₂ rises, driving climate change. Human pressure on food webs — overharvesting top consumers, simplifying communities — also disrupts energy flow through ecosystems. At this level you need only the outline: human activity adds CO₂ to the atmosphere faster than the carbon cycle can remove it, unbalancing a cycle that was previously in near-equilibrium.
Common mistakes examiners penalise
Saying energy is 'recycled' — energy FLOWS through and is lost as heat; only MATTER cycles. This is the single most penalised confusion in the topic.
Drawing food-chain arrows backwards — arrows point in the direction energy flows, from prey to predator, not 'who eats whom'.
Explaining the 90% loss vaguely — you must name the losses: heat from RESPIRATION (the main one), material not eaten, and material not digested (lost in faeces). 'Energy is used up' scores nothing.
Confusing producers and decomposers — producers MAKE organic matter (photosynthesis); decomposers BREAK it DOWN and release nutrients. They are opposite roles.
Treating the 10% rule as an exact law — it is an approximation (about 5–20%); state it as a rule of thumb, not a fixed figure.
Saying a pyramid of energy 'can be inverted' — pyramids of numbers and biomass can be, but a pyramid of energy is ALWAYS upright because energy is lost at every step.
Muddling carbon-cycle processes — photosynthesis REMOVES CO₂; respiration, decomposition and combustion RETURN it. Don't assign a process to the wrong direction.
Forgetting decomposers in nutrient cycling — nutrients cannot return to producers without decomposition; omitting decomposers leaves the cycle broken.
Model answer — marked the way our engine marks it
This is an EXPLAIN question, and explain marks are awarded analytically — each distinct valid biological point is worth one mark, up to the number of marks available. There is no need for a fixed wording: the engine looks for separate creditable ideas, awards the answer mark (A) for a correctly reasoned conclusion, applies error-carried-forward (ECF) so an earlier slip does not sink later points, and accepts equivalent phrasings. Study how each mark below is tied to a specific, named idea.
Where this leads
The two rules of this topic — energy flows, matter cycles — underpin the rest of ecology. The ten-percent ceiling explains why eating lower on a food chain feeds more people from the same land, and why top predators are always few. Nutrient cycling links straight to the nitrogen and phosphorus cycles and to conservation, while the carbon cycle connects your biology to the physics of the greenhouse effect and to the whole question of human-driven climate change. Hold the core distinction firmly — energy in as light and out as heat, atoms round and round forever — and the rest of ecosystem ecology falls into place.
Worked examples
See the formulas applied — reveal one step at a time, like the exam.
A meadow's producers store 40 000 kJ m⁻² yr⁻¹ of energy as new biomass. The primary consumers that feed on them store 3200 kJ m⁻² yr⁻¹. (a) Calculate the percentage efficiency of energy transfer from producers to primary consumers. (b) State whether this is consistent with the 10% rule, and explain in one sentence where most of the 'missing' energy has gone. [4]
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(a) Percentage efficiency. Efficiency (%) = (energy stored at the higher level ÷ energy available at the lower level) × 100. [M1: correct method] Efficiency = (3200 ÷ 40 000) × 100 = 8.0%. [A1: 8.0%]
The producers in a pond fix 50 000 kJ m⁻² yr⁻¹ of energy. Assuming exactly 10% is transferred at each step, estimate the energy available to the tertiary consumers, and use your answer to explain why the pond cannot support a fifth trophic level. [4]
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Step 1 — apply the 10% rule level by level. Each arrow passes on one tenth of the energy.
- Producers: 50 000 kJ m⁻² yr⁻¹.
- Primary consumers: 10% of 50 000 = 5000 kJ m⁻² yr⁻¹. [M1: 10% applied correctly]
- Secondary consumers: 10% of 5000 = 500 kJ m⁻² yr⁻¹.
- Tertiary consumers: 10% of 500 = 50 kJ m⁻² yr⁻¹. [A1: 50 kJ m⁻² yr⁻¹]
Explain why the amount of energy available decreases at each successive trophic level in a food chain. [4]
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Model answer. At each trophic level, organisms lose much of their energy as heat through respiration, because they respire to power life processes such as movement and growth. In addition, not all of an organism at one level is eaten by the next, and of the material that is eaten, not all is digested and absorbed — the indigestible part is lost in faeces. As a result, only about 10% of the energy at one level is stored as new biomass and so passed on to the next. Because roughly 90% is lost at every transfer, the energy available falls sharply at each successive level, which is also why food chains are limited in length.
How it all connects
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Glossary
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Quick check
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Revision flashcards
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Open system (ecosystem)
A system that exchanges both energy AND matter with its surroundings. Ecosystems are open: energy enters as light and leaves as heat, and matter can enter and leave — but crucially, energy FLOWS through while matter CYCLES within.
Key takeaways
Review these before you close the topic — retrieval beats re-reading.
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Energy FLOWS: one direction only — in as light, out as heat; lost at every step; never recycled; needs constant resupply from the Sun.
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Matter CYCLES: the same finite atoms are used again and again, passing between organisms and the environment.
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Open system: an ecosystem exchanges both energy and matter with its surroundings.
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The commonest error in this whole topic is saying energy is 'recycled' — it is not.
Practice — then mark it
The whole point: a real Cambridge question, marked mark-by-mark.
Get a Paper 2 question marked: explain why energy decreases along a food chain, or work through a carbon-cycle / efficiency question with full reasoning
Get a Paper 2 question marked: explain why energy decreases along a food chain, or work through a carbon-cycle / efficiency question with full reasoning
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Checkpoint
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Before you move on: do Get a Paper 2 question marked: explain why energy decreases along a food chain, or work through a carbon-cycle / efficiency question with full reasoning on paper, snap a photo, and get examiner-style feedback on exactly where you win and lose marks.