In simple terms
A friendly intro before the formal notes — no formulas yet.
The Ecosystem's Economy
Ecosystems run on an energy budget, where energy flows through once and is lost, while essential nutrients are constantly recycled. Productivity measures how efficiently organisms convert energy into biomass.
Think of an ecosystem's energy flow like your personal finances. Your gross salary is the total energy captured (Gross Productivity). After you pay for your essential living costs like rent and food (Respiration), the money you have left to save or spend on growth is your net income (Net Productivity). You can't recycle the money you've spent, it's gone, just like energy is lost as heat.
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First, identify the source of energy (usually the sun) and trace its path through different trophic levels, remembering that energy is lost at each step.
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Next, distinguish between Gross Productivity (total energy assimilated) and Net Productivity (energy left for growth after respiration). Use the formulas NPP = GPP - R and NSP = GSP - R.
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Then, analyse nutrient cycles like carbon and nitrogen by identifying the main storages (sinks) and the processes that move nutrients between them (flows or fluxes).
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Finally, evaluate human impacts on these cycles, such as burning fossil fuels (carbon cycle) or using fertilisers (nitrogen cycle), and their consequences.
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Key formulas
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Full topic notes
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Energy Flow and the Laws of Thermodynamics
Energy in almost all ecosystems originates from the sun. This solar energy is captured by autotrophs (producers) and converted into chemical energy through photosynthesis. This energy is then transferred through the ecosystem as one organism consumes another. However, this flow is unidirectional and inefficient.
First Law of Thermodynamics: Energy can be transferred and transformed, but it cannot be created or destroyed. In ecosystems, this means solar energy is converted to chemical energy, which is then transferred between trophic levels.
Second Law of Thermodynamics: During any energy conversion, some energy is lost as heat, and the entropy (disorder) of the universe increases. This explains why energy transfers are inefficient (the '10% rule') and why food chains are limited in length. Energy is lost as heat during respiration at each trophic level.
Ecosystem Productivity
Productivity is the rate at which biomass is generated in an ecosystem. It is a key measure of an ecosystem's health and ability to support life. We distinguish between primary productivity (by producers) and secondary productivity (by consumers).
For Producers (Autotrophs):<br><b>Net Primary Productivity (NPP) = Gross Primary Productivity (GPP) - Respiration (R)</b><br><br>GPP is the total energy fixed by photosynthesis. R is the energy used by the producers for their own life processes. NPP is the energy available to herbivores.
For Consumers (Heterotrophs):<br><b>Net Secondary Productivity (NSP) = Gross Secondary Productivity (GSP) - Respiration (R)</b><br><br>GSP is the energy assimilated from food (). R is the energy used by the consumer for its life processes. NSP is the formation of new biomass (growth and reproduction).
Nutrient Cycling: Carbon and Nitrogen
Unlike energy, which flows through an ecosystem and is lost, matter is cycled. Biogeochemical cycles describe the pathways of essential elements like carbon and nitrogen through the biotic (living) and abiotic (non-living) components of an ecosystem. We model these using storages (where the nutrient is held) and flows (the processes that move it).
Carbon Cycle: Key storages include the atmosphere (), oceans, biomass, and fossil fuels. Key flows are photosynthesis (atmosphere to biomass), respiration (biomass to atmosphere), combustion (fossil fuels to atmosphere), and diffusion (atmosphere to ocean).
Nitrogen Cycle: The main storage is the atmosphere (). Key flows include nitrogen fixation (by bacteria or lightning, converting to ammonia), nitrification (ammonia to nitrates), assimilation (uptake by plants), denitrification (nitrates to ), and ammonification (decomposition).
Human Impact: Burning fossil fuels adds excess to the atmosphere, enhancing the greenhouse effect. The industrial production of fertilisers (Haber-Bosch process) has doubled the rate of nitrogen fixation, leading to eutrophication in aquatic systems.
Worked examples
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In a temperate forest ecosystem, the Gross Primary Productivity (GPP) was measured to be 8,400 . The plant community's respiratory losses (R) were found to be 4,600 . Calculate the Net Primary Productivity (NPP) of this ecosystem.
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<b>1. State the formula:</b><br>NPP = GPP - R [1 mark]<br><br><b>2. Substitute the values:</b><br>NPP = 8,400 - 4,600 [1 mark for correct substitution]<br><br><b>3. Calculate the final answer with units:</b><br>NPP = 3,800 [1 mark for correct answer and units]<br><br><b>Total: 3 marks</b>
A population of rabbits in a meadow consumes 25,000 of plant matter. They produce 16,000 of faeces (egestion). Their respiratory heat loss is 7,500 . a) Calculate the Gross Secondary Productivity (GSP). b) Calculate the Net Secondary Productivity (NSP).
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<b>a) Calculate GSP:</b><br><b>1. State the formula:</b> GSP = Food eaten - Faecal loss [1 mark]<br><b>2. Substitute values:</b> GSP = 25,000 - 16,000 <br><b>3. Final Answer:</b> GSP = 9,000 [1 mark]<br><br><b>b) Calculate NSP:</b><br><b>1. State the formula:</b> NSP = GSP - R [1 mark]<br><b>2. Substitute values:</b> NSP = 9,000 - 7,500 <br><b>3. Final Answer:</b> NSP = 1,500 [1 mark]<br><br><b>Total: 4 marks</b>
How it all connects
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Glossary
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Gross Primary Productivity (GPP)
The total amount of chemical energy produced by autotrophs in an ecosystem, typically through photosynthesis, per unit area per unit time. It's the total 'income' of energy before any is used. Units: or .
Key takeaways
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First Law of Thermodynamics: Energy can be transferred and transformed, but it cannot be created or destroyed. In ecosystems, this means solar energy is converted to chemical energy, which is then transferred between trophic levels.
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Second Law of Thermodynamics: During any energy conversion, some energy is lost as heat, and the entropy (disorder) of the universe increases. This explains why energy transfers are inefficient (the '10% rule') and why food chains are limited in length. Energy is lost as heat during respiration at each trophic level.
Practice — then mark it
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Test Your Knowledge on Energy Flow and Nutrient Cycles
Test Your Knowledge on Energy Flow and Nutrient Cycles
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